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Raichlen, D., Gordon, A., Foster, A., Webber, J., Sukhdeo, S., Scott, R., Gosman, J., & Ryan, T. M. (in press) An ontogenetic framework linking locomotion and trabecular bone architecture with applications for reconstructing hominin life history. Journal of Human Evolution.

Ryan, T. M. & Sukhdeo, S. M. (in press) KSD-VP 1/1 Analysis of the Postcranial Skeleton Using High-Resolution Computed Tomography. In Geology, Chronology, and Anatomy of KSD-VP-1/1, Woranso-Mille, Ethiopia.  (Y. Haile-Selassie and Denise Su, eds). Springer Publishing.

Ryan, T. M. & Shaw, C. N. (2015) Gracility of the modern Homo sapiens skeleton is the result of decreased biomechanical loadingProceedings of the National Academy of Sciences, USA. 112:372-377.

Hung, N. M., Ryan, T. M., Stauffer, J. R., & Madsen, H. (2015) Does hardness of food affect the development of pharyngeal teeth of the black carp, Mylopharyngodon piceus (Pisces: Cyprinidae)? Biological Control 80:156-159.



Shaw, C. N., Stock, J.T., Davies, T. G., & Ryan, T. M. (2014) Does the distribution and variation in cortical bone along lower limb diaphyses reflect selection for locomotor economy? In Mobility: Towards a Definition for Application in Human Evolution. (D. Marchi and K. Carlson, eds) Springer Publishing. pp. 49-66.



Ryan, T. M., & Shaw, C. N. (2013). Trabecular bone microstructure scales allometrically in the primate humerus and femur. Proceedings of the Royal Society B: Biological Sciences 280:20130172.

Gosman, J. H., Hubbell, Z. R., Shaw, C. N., & Ryan, T. M. (2013) Development of cortical bone geometry in the human femoral and tibial diaphysis. Anatomical Record 296:774-787.

Macintosh, A. A., Davies, T. G., Ryan, T. M., Shaw, C. N., & Stock, J. T. (2013). Periosteal versus true cross-sectional geometry: A comparison along humeral, femoral, and tibial diaphyses. American Journal of Physical Anthropology 150:442-452.



Ryan, T. M., & Shaw, C. N. (2012). Unique suites of trabecular bone features characterize locomotor behavior in human and non-human anthropoid primates. PLoS ONE, 7(7):e41037, 11.

Ryan, T. M., Silcox, M. T., Walker, A., Mao, X., Begun, D. R., Benefit, B. R., Gingerich, P. D., Kohler, M., Kordos, L., McCrossin, M. L., Moya-Sola, S., Sanders, W. J., Seiffert, E. R., Simons, E., Zalmout, I. S., & Spoor, F. (2012). Evolution of locomotion in Anthropoidea: the semicircular canal evidence. Proceedings of the Royal Society B: Biological Sciences 279:3467–3475.

Shaw, C. N., & Ryan, T. M. (2012). Does skeletal anatomy reflect adaptation to locomotor patterns? Cortical and trabecular architecture in human and non-human anthropoids. American Journal of Physical Anthropology 147:187-200.



Dumont, E. R., Ryan, T. M., & Godfrey, L. R. (2011). The Hadropithecus conundrum reconsidered, with implications for interpreting diet in fossil hominins. Proceedings of the Royal Society B 278:3654–3661.

Vinyard, C. J., Taylor, A. B., Teaford, M. F., Glander, K.E., Ravosa, M. J., Rossie, J. B., Ryan, T. M., & Williams, S. H. (2011). Are we looking for loads in all the right places? New research directions for studying the masticatory apparatus of New World monkeys. Anatomical Record 294:2140-2157.

Hogg, R., Vinyard, C. J., Ryan, T. M., & Ravosa, M. J. (2011). The functional morphology of the anterior masticatory apparatus in tree-gouging marmosets (Cebidae, Primates). Journal of Morphology 272:833-849.



Seiffert, E. R., Simons, E. L., Boyer, D. M., Perry, J. M., Ryan, T. M., & Sallam, H. M. (2010). A peculiar primate of uncertain affinities from the earliest late Eocene of Egypt. Proceedings of the National Academy of Sciences of the USA 107:9712-9717.

Martínez-Abadías, N., Percival, C., Aldridge, K., Hill, C. A., Ryan, T. M., Sirivunnabood, S., Wang, Y., Jabs, E. W., & Richtsmeier, J. T. (2010). Beyond the closed suture in apert syndrome mouse models: Evidence of primary effects of FGFR2 signaling on facial shape at birth. Developmental Dynamics 239:3058-3071.

Griffin, N. L., D’Août, K., Ryan, T. M., Richmond, B. G., Ketcham, R. A., & Postnov, A. (2010). Comparative forefoot trabecular bone architecture in extant hominids. Journal of Human Evolution 59:202-213.

Ryan, T. M., M. Colbert, R.A. Ketcham, and C.J. Vinyard.  (2010) Trabecular bone structure in the mandibular condyle of gouging and non-gouging platyrrhines. American Journal of Physical Anthropology 141:583–593.

Ryan, T. M. and A. Walker (2010) Trabecular bone structure in the humeral and femoral heads of anthropoid primates. Anatomical Record 293:719–729.



Silcox, M.T., J.I. Bloch, D.M. Boyer, M. Godinot, T. M. Ryan, F. Spoor, and A. Walker. (2009) The semicircular canal system in early primates and euprimates. Journal of Human Evolution 56:315-327.

Welker, K., J. Orkin, and T. M. Ryan. (2009) Analysis of intra-individual and inter-sex variation in semi-circular canal dimensions using high-resolution x-ray computed tomography. Journal of Anatomy 215:444-451.



Ryan, T.M., D.A. Burney, L.R. Godfrey, U.B. Göhlich, W.L. Jungers, N. Vasey, Ramilisonina, A. Walker, G.W. Weber. (2008) A reconstruction of the Vienna skull of Hadropithecus stenognathus. Proceedings of the National Academy of Sciences. USA 105:10699-10702.

Jeffery, N., T. M. Ryan and F. Spoor. (2008) The primate subarcuate fossa and its relationship to the semicircular canals part II: adult interspecific variations. Journal of Human Evolution 55:326-339.

Walker, A., T. M. Ryan, M.T. Silcox, E. Simons, F. Spoor. (2008) The semicircular canal system and locomotion: the case of extinct lemuroids and lorisoids. Evolutionary Anthropology 17:135-145.



Seiffert, E.R., E.L. Simons, T. M. Ryan, T.M. Bown, and Y. Attia. (2007) New remains of Eocene and Oligocene Afrosoricida (Afrotheria) from Egypt, with implications for the origin(s) of afrosoricid zalambdodonty. Journal of Vertebrate Paleontology 27:963-972.

Spoor, F., T. Garland Jr., G. Krovitz, T. M. Ryan, M.T. Silcox, and A. Walker. (2007) The primate semicircular canal system and locomotion. Proceedings of the National Academy of Sciences USA 104:10808-10812.

Fraser, N.E., P. E. Olsen, A.C. Dooley, and T.M. Ryan. (2007)   A new gliding tetrapod (Diapsida: ?Archosauromorpha) from the Upper Triassic (Carnian) of Virginia. Journal of Vertebrate Paleontology 27:261-265.

Simons, E.L., E.R. Seiffert, T.M. Ryan, and Y. Attia. (2007) A remarkable female cranium of the early Oligocene anthropoid Aegyptopithecus zeuxis (Catarrhini, Propliopithecidae). Proceedings of the National Academy of Sciences USA 104:8731-8736.

Parsons, T., T. M. Ryan, R.H. Reeves, and J. Richtsmeier. (2007) Micro-structure of trabecular bone in a mouse model for Down syndrome. The Anatomical Record 290:414-421.



Vinyard, C. J. and T. M. Ryan. (2006) Cross-sectional bone distribution in the mandibles of gouging and non-gouging platyrrhines.International Journal of Primatology 27:1461-1490.

Ryan, T. M. and G.E. Krovitz. (2006) Trabecular bone ontogeny in the human proximal femur. Journal of Human Evolution 51:591-602.

Ryan, T.M. and G.R. Milner. (2006) Osteological applications of high-resolution computed tomography: A prehistoric arrow injury.Journal of Archaeological Science 33:871-879.

Maga, M., J. Kappelman, T.M. Ryan, and R.A. Ketcham. (2006) The calcaneal trabecular microarchitecture of extant large bodied hominoids. American Journal of Physical Anthropology 129:410-417.



Seiffert, E.R., E.L. Simons, T.M. Ryan, and Y. Attia. (2005) Additional remains of Wadilemur elegans, a primitive stem galagid from the late Eocene of Egypt. Proceedings of the National Academy of Sciences USA. 102:11396-11401.

Ryan, T. M. and R. A. Ketcham. (2005) The angular orientation of trabecular bone in the femoral head and its relationship to hip joint loads in leaping primates. Journal of Morphology 265:249-263.

Ryan, T. M. and B. van Rietbergen. (2005) Mechanical significance of femoral head trabecular bone structure in Loris and Galago evaluated using micromechanical finite element models. American Journal of Physical Anthropology 126:82-96.



Ketcham, R. A. and T. M. Ryan. (2004) Quantification of anisotropy in trabecular bone. Journal of Microscopy 213:158-171.



Kappelman, J., D. T. Rasmussen, W. J. Sanders, M. Feseha, T. Bown, P. Copeland, J. Crabaugh, J. Fleagle, M. Glantz, A. Gordon, B. Jacobs, M. Maga, K. Muldoon, A. Pan, L. Pyne, B. Richmond, T. Ryan, E. R. Seiffert, S. Sen, L. Todd, M. C. Wiemann, A. Winkler. (2003) Oligocene mammals from Ethiopia and faunal exchange between Afro-Arabia and Eurasia. Nature 426:549-552



Ryan, T. M. and R. A. Ketcham. (2002) Femoral head trabecular bone structure in two omomyid primates. Journal of Human Evolution 43:241-263.

Ryan, T. M. and R. A. Ketcham. (2002) The three-dimensional structure of trabecular bone in the femoral head of strepsirrhine primates. Journal of Human Evolution 43:1-26.

Fajardo, R. J., T. M. Ryan, and J. Kappelman. (2002) Assessing the accuracy of high-resolution X-ray computed tomography of primate trabecular bone by comparisons with histological sections. American Journal of Physical Anthropology 118:1-10.